The causes the lips to close and pucker during kissing, whistling and speaking.

The Orbicularis oris causes the lips to close and pucker during kissing, whistling and speaking.

Explanation:

Orbicularis oris muscles are facial muscles which are located in the thickness of lip around oral opening.

The muscle consists of two main parts

(a) Marginal part consisting of the longitudinal fibers -Closes the oral opening.

(b)Labial part consisting of the circular fibers - pushes the lips forward.

The main function of the orbicularis oris is closing of the mouth.  It puckers the lips when it contracts.

These muscles are used to do actions like kissing, whistling and speaking.

Thus,

The Orbicularis oris causes the lips to close and pucker during kissing, whistling and speaking.

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