The glycerol-3-phosphate shuttle can transport cytosolic NADH equivalents into the mitochondrial matrix (see Fig. 15.11c). In this shuttle, the protons and electrons are donated to FAD, which is reduced to FADH2. These protons and electrons are subsequently donated to coenzyme Q in the electron transport chain. Given that the number of ATP molecules made per NADH and FADH2 oxidation differ by ? the amount of ATP generated per mole of glucose when the glycerol-3-phosphate shuttle would be instead of 32.