What determines which base is to be added to an rna strand during transcription?

sheep is a herbivore animal whereas grasshopper belongs to the class chordates but both of them feed on greens.  

if grasshopper and sheep feeding on a pasture land, i do not think both of them will have any impact on each other. however, the number of the species can make difference. since sheep feed on greens, there is a possibility there left no pasture land for the grasshoppers or they may get extinct.

1. is geographic distribution

2 is increasing complexity

3 is transitional forms